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A close-up of a Devonshire cup coral attached to a raised cave shelf. Diamond Cave, Fair Isle. ©Graham Saunders

A close-up of a Devonshire cup coral attached to a raised cave shelf. Diamond Cave, Fair Isle. ©Graham Saunders

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Dead man's fingers nestling amongst a dense turf of oaten pipes hydroids on the upper cave wall. Diamond Cave, Fair Isle. ©Graham Saunders

Dead man's fingers nestling amongst a dense turf of oaten pipes hydroids on the upper cave wall. Diamond Cave, Fair Isle. ©Graham Saunders

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Starfish hanging on to a mat of colonial baked-bean sea squirts, commonly found on the upper cave walls. Diamond Cave, Fair Isle. ©Graham Saunders

Starfish hanging on to a mat of colonial baked-bean sea squirts, commonly found on the upper cave walls. Diamond Cave, Fair Isle. ©Graham Saunders

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The delicate white lace sponge and baked-bean sea squirts. This sponge sometimes covers considerable areas of the cave walls. Diamond Cave, Fair Isle. ©Graham Saunders

The delicate white lace sponge and baked-bean sea squirts. This sponge sometimes covers considerable areas of the cave walls. Diamond Cave, Fair Isle. ©Graham Saunders

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The beadlet anemone is one of the few animals that are able to survive in the violent surge conditions of the cave floor. South Haven Cave, Fair Isle. ©Graham Saunders

The beadlet anemone is one of the few animals that are able to survive in the violent surge conditions of the cave floor. South Haven Cave, Fair Isle. ©Graham Saunders

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A squat lobster and a closed beadlet anemone on a typical boulder cave floor. Diamond Cave, Fair Isle. ©Graham Saunders

A squat lobster and a closed beadlet anemone on a typical boulder cave floor. Diamond Cave, Fair Isle. ©Graham Saunders

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A small diamond-shaped hole in the roof of the cave gives the cave its name and shines a dim light on the dense clumps of oaten pipes hydroids that colonise the larger boulders. ©Graham Saunders

A small diamond-shaped hole in the roof of the cave gives the cave its name and shines a dim light on the dense clumps of oaten pipes hydroids that colonise the larger boulders. ©Graham Saunders

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Vertical cave wall with dead man's fingers and urchin with a surrounding species-rich turf of hydroids, sea squirts and sponges. Diamond Cave, Fair Isle. ©Graham Saunders

Vertical cave wall with dead man's fingers and urchin with a surrounding species-rich turf of hydroids, sea squirts and sponges. Diamond Cave, Fair Isle. ©Graham Saunders

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Because of the potentially inhospitable and hazardous environment, an experienced cave diver supplemented the biological survey team, providing additional safety and line-laying support. ©Graham Saunders

Because of the potentially inhospitable and hazardous environment, an experienced cave diver supplemented the biological survey team, providing additional safety and line-laying support. ©Graham Saunders

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Arctic Skua, Isle of Noss NNR, Shetland. ©Lorne Gill/NatureScot

Arctic Skua, Isle of Noss NNR, Shetland. ©Lorne Gill/NatureScot

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